In TowerFall: Ascension, Sacred Orbs are unique collectibles that players encounter during their runs in Quest mode. While the specifics of their gameplay mechanics aren’t exhaustively detailed in the available literature, some educated assumptions can be drawn based on general game design principles and the nature of similar collectibles found in the game.

Quest Mode Overview
Quest mode in TowerFall: Ascension tasks players with navigating through various stages where they must eliminate waves of enemies. Players have limited lives and must perform well under either Normal or Hardcore difficulty settings. Successfully completing these stages not only requires strategic gameplay but also includes the challenge of optimizing performance within a limited time frame.

Sacred Orbs
While direct information on Sacred Orbs is scarce, it is worth considering the typical roles of collectibles in similar gaming contexts:

1. Power-ups and Bonuses: Sacred Orbs likely offer players various enhancements or bonuses that can aid in combat, increase survivability, or enhance abilities during their runs. This could range from temporary invincibility, increased damage, or additional arrows.

2. Completion Rewards: Such items often contribute to the completion percentage of a stage or unlock additional content, such as new characters, stages, or game modes upon successful collection.

3. Score Boosts: Given that TowerFall rewards players for performance, collecting Sacred Orbs might also provide a score boost, influencing the final evaluation of the player’s run.

4. Unlocks and Progression: Collectibles in games usually contribute to broader progression systems. Sacred Orbs may be linked to achievements or milestones that, when accumulated, could unlock new features or lore within the game.

Conclusion
Without definitive data on the specifics of Sacred Orbs in Quest mode, it remains a challenge to provide precise information. However, they presumably play a role similar to other collectibles in enhancing gameplay or unlocking new content. For detailed mechanics, players may benefit from engaging with community resources, including fan websites, forums, or even in-game archives, which can offer deeper insights. Exploring these avenues might yield the comprehensive information necessary to fully understand the impact of Sacred Orbs in TowerFall: Ascension’s Quest mode.